WCA-3A, and ENP-TS sites. However, these spikes were not correlated with increased

CRR.

 The significant benthic BGL and LEU increase at the WCA-2A transitional site is

accompanied by a generally much lower TN (31 g kg-1), lignin (21%), cellulose (9%), the

lowest TOC (352 g kg-1) of any site not in ENP-TS, as well as elevated Ca (81 g kg-1) and

Mg (81 mg kg-1 ). This site is also dominated by Chara spp., which is a characteristic

algae in transitional P sites and is a level II indicator of P enrichment (U.S. EPA, 2002).

The spike in the benthic LNWR transitional site is also accompanied by a lower TN

(31.23 g kg-1) and TOC (438.1 g kg-1).

 Increases also occurred in the soil LEU and PHO at the transitional sites in WCA-

2A and ENP-TS. The lowest TP (0.2 g kg-1) and TN (6 g kg-1) of any site was recorded

at the transitional site in ENP-TS which accounts for the greatest PHO and LEU activity

of any soil layer. This relationship in the soil layer is not as clear at the WCA-2A

transitional site.

 Discussion

 The following discussion has been grouped into several sections that first

compare the general effects of the P gradient between the enriched and reference sites.

The discussion generally pertains to the benthic layer results, as this layer is the most

biologically active (White and Reddy, 2000) and is more responsive to changes in

environmental conditions. Secondly, specific hydrologic units are compared in relation

to their response to the P gradient. Lastly, a brief discussion of the observations made at

the transitional sites is included.