Table 4-2.


Benthic nutrient, enzyme, and ratio parameters in P enriched and reference sites in WCA-3A and ENP-TS. Values
presented are means averaged over the three sampling periods with standard errors. Significant differences are given
between the enriched and background sites in the respective areas. Each asterisk reflects differences present at the
p<0.05 level per sampling period. Analysis was divided into the three sampling periods (1, 2, 3). Glucosidase, Leucine
aminopeptidase and Phosphatase units are moles AMC-MUF released g-1 AFDM h-1. Phenol oxidase and Peroxidase
units are moles DICQ released g-1 AFDM h-1.


 WCA-3A ENP-TS

Parameter Impacted Reference (P<0.05) Impacted Reference (P<0.05)
 (n=9) (n=9) 1 2 3 (n=9) (n=9) 1 2 3


Total phosphorus
Total nitrogen
Total organic carbon
Calcium
Magnesium
C:P
C:N
N:P
Lignin
Cellulose
Glucosidase
Leucine aminopeptidase
Phosphatase
Phenol oxidase
Peroxidase
Ecell/Ep
Ecell/En
Ecell/Eox


g kg-1
g kg-1
g kg-1
g kg-1
g kg-1



%


1.66 0.25
33.16 1.12
438.11 9.99
64.72 39.35
1.17 0.10
283.7 48.8
13.2 0.3
21.6 4.3
21.6 4.0
15.4 4.7
1.07 0.29
4.39 0.65
2.32 0.56
186.6 46.0
10.8 4.3
53.20 22.73
1.33 0.14
4.22 1.01


0.48 0.05
37.53 1.95
427.67 16.88
25.78 6.34
1.13 0.16
840.8 5.4
11.5 0.3
75.2 0.4
26.5 2.7
11.5 0.4
0.26 0.02
3.23 0.10
18.09 3.23
105.2 4.1
17.3 4.1
1.51 0.61
0.57 0.09
1.44 0.46


* 0.74 0.20
 13.2 4.77
 173.63 46.82
 166.44 58.77
 1.43 0.05
* 236.3 21.6
* 13.1 + 1.5
* 18.2 1.1
 20.1 + 11.1
 20.9 4.8
 0.67 0.01
 8.13 0.79
* 11.19 3.18
 287.3 89.1
 10.8 2.2
* 3.47 0.80
 1.59 0.41
 2.22 0.43


0.10 + 0.01
11.98 0.90
168.00 + 8.19
191.11 28.04
2.44 0.16
1803.2 137.9
14.4 0.1
123.6 8.3
7.2 4.5
8.4 5.0
0.15 0.01
3.07 0.05
61.33 0.48
94.9 2.3
10.4 2.2
0.11 0.02
0.44 0.09
0.96 0.20


*1 *