pdu operon pocR pduF AB C D E G H J K L M N 0 P Q S T U VW X I I I I I I 1 ? ? 1 1 I I 1 ? 1 1 I ? 1 I ? 1 D Carboxysome protein homolog D Enzyme E-l Unknown Figure 1-3. The pdu operon. The genes of the pdu operon are divided into three groups indicated by the shading of the boxes. The functions of the pduABCDEGHJKO and pduP genes have been assigned based on homology to known proteins as well as the results of biochemical and genetic studies. The functions of the remaining proteins are based on homology alone.