THOMPSON: Humboldtiana from Chihuahua and Texas
97
Remarks. The holotype is re-figured to show all faces of the shell (Figs 78-81).
ACKNOWLEDGMENTS
I am grateful to the following people who have assisted me in this study. John Anderson, (Houston Texas, USA), Richard L. Thompson and Daniel Thompson (Stafford, Texas, USA), provided field assistance at the Coal Mine Ranch, Presidio Co., Texas. Jon Vogler (Houston; Texas, USA) and Fred Nelan gave permission to visit the Coal Mine Ranch and the generously hosted us during our visit. Elizabeth Mihalcik (Bainbridge, Georgia, USA), provided field assistance in Chihuahua, Mexico. I wish to thank Jonathan Ablett (BMNH), Dan L. Graf (ANSP) and Robert Hershler (USNM) for the loan of specimens in their charges. Charlotte M. Porter (Florida Museum of Natural History) assisted with derivations of Classical Greek nomenclature.
LITERATURE CITED
Burch, J. B. & F. G. Thompson. 1957. Three new Mexican land snails of the genus Humboldtiana. Occasional Papers Museum of Zoology, University of Michigan, (590): 1-11.
Fullington, R. W. & E. L. Zimmerman. 1977. A new species of Humboldtiana (Helminthoglyptidae) from Coahuila, Mexico. Veliger,20: 134-136.
Martens, E. von. 1890-1901. Biologia Centrali-Ameri-cana. Mollusca. 1-706. British Museum (Natural History).
Metcalf, A. L. 1983. Anew Humboldtiana (Puimonata; Helminthoglytidae) from northwestern Coahuila, Mexico. Nautilus, 97: 69-72.
Metcalf, A. L. 1984. A new Humboldtiana (Pulmonata: Helminthoglyptidae) from extreme eastern Chihuahua, Mexico. Nautilus, 98: 145-147.
Pilsbry, H. A. 1927. The structure and affinities of Humboldtiana and related helicid genera of Mexico and Texas. Proceedings of the Academy of Natural Science of Philadelphia, 79: 165-192.
Pilsbry, H. A. 1935a. Western and southwestern Amnicolidae and a new Humboldtiana. Nautilus^: 91-94.
Pilsbry, H. A. 1935b. Descriptions of Middle American Land and freshwater Mollusca. Proceedings of the Academy of Natural Science of Phila delphia 87:1-6.
Pilsbry, H. A. 1939. Land Mollusca of North America. Vol. l,Pt. 1. Philadelphia Academy of Natural Sciences. 395-410.
the embryonic whorls and the absence of granular sculpture on the postembryonic whorls immediately distinguish it from all other groups within the genus. The species is variable in relative height as well as in the color pattern. Two paratypes (Figs. 74-75) are illustrated for comparison with the holotype (Figs. 70-73).
Etymology. The name tanymastix (f.) is from the Classical Greek tocvu, meaning long and paaxi^, a whip, alluding to the long flagellum on the epiphallus.
Undetermined subgenus
The following two species from Chihuahua are referred to Humboldtiana bicincta species group {see Thompson & Brewer, 2000). Both are unknown anatomically, so their placement in this group is provisional. They have a depressed spire that is about 0.7 times the shell width, they have three bands on a white background, and they have smooth shells that lack granular sculpture. In some species of this group the bands may be obsolete or entirely absent.
Humboldtiana eulaliae Metcalf, 1984
Humboldtiana eulaliae Metcalf, 1984; Nautilus, 98:145-147; figs. 1-4 (shell).
Type locality. Chihuahua, west side of the Sierra Santa Eulalia (2712.0'N, 10347.6'W); 1250 m alt. Holotype USNM 820297; collected by Wally Lippencott (U. S. Department of Agriculture), 23 March, 1982.
Metcalf provides the following information about the type locality ... "is along walls of the canyon debauching southwestward about midway of the Santa Eulalia range. The mouth of the canyon is 1.3 km E of El Pinalero and 7.5 km N and 1.5 km E of El "Penoles" on the Guimbalete quadrangle. The canyon is ca. 2.5 km long heading at 1650 m and debauching at 1250 m".
Distribution. Known only from the type locality.
Humboldtiana torrei Pilsbry, 1935
Humboldtiana torrei Pilsbry, 1935b: 1; 2; pi. 1, fig. 10 (shell).
Type locality. Chihuahua, San Antonio y Santa Rosalia, near and south of Cd. Chihuahua. Holotype: ANSP 162325; collected by Carlos de la Torre, about 1899. Paratypes ANSP (1), Carlos de la Torre collection, Havana, Cuba.
Distribution. Known only from the type locality.