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BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 46(3)
tions interrupted by occasional white streaks. The last whorl descends strongly to the aperture to insert at the bottom of the sub-peripheral band. The plane of the aperture lies at about 40 to the shell axis in lateral profile (Fig. 37). The aperture is circular, 0.66-0.80 times as wide as high. The outer lip and the basal lip are very slightly reflected. The columellar lip is widely reflected over the umbilicus. The parietal wall of the aperture has a thin transparent glaze.
Measurements of the holotype and six paratypes are given in Table 5.
Anatomy (Figs. 34-35). (Paratypes, UF 358375; 4 specimens). The specimens have been preserved for 35 years in 70% ethanol. They are light gray on the head-foot, mantle collar and lung.
The albumen gland is elongate and stout. The carrefour is exposed. The genital atrium is very short, and is separated from the body wall by 1-2 mm. The penis is short, stout and nearly uniform-cylindrical in shape externally. It is 7 mm long, and is demarcated from the epiphallus by a weak constriction. Internally the penis wall is lined with heavy longitudinal pilasters (Fig. 35). The upper part of the chamber bears a short four-lobed verge. The penis retractor muscle is 22-23 mm long, slender. It originates on the middle of the inner wall of lung behind the mantle collar, and inserts on and forms a sheath around the lower third of the epiphallus. The epiphallus is very long, stout and thick-walled. It is about 5-6 times as long as the penis, and it is lined internally with four longitudinal glandular columns. The flagellum is short and stout. It is slightly shorter than the epiphallus and is lined internally with six longitudinal columns of papillae. The vas deferens is moderately stout and is nearly uniform in diameter throughout its length. The lower vagina is moderately long, slender and conical with four unequal-sized dart-sacs radially arranged above. One-dart sac, ds, is very large; the other three dart-sacs are much smaller and are sub-equal to each other in size. A stout dart-gland encircles the vagina immediately above, but not appressed
against the dart-sacs. The free vagina is almost absent. The spermathecal duct diverges from the vagina just above the dart-glands; it is relatively stout, very long, and bears a short caecum at 0.69-0.75 of the distance from the base. The caecum is about a third or less than the length of the distal spermathecal duct segment. The spermatheca is small and elongate-bulbous in shape. It is oppressed against the upper end of the prostate-uterus at the base of the albumen gland. The spermatheca duct + spermatheca is about twice the length of the uterus.
Type locality. Chihuahua, a rocky outcrop in the foothills of the Sierra La Catarina, Buenaventura, (2940'N, 10730'W); 1850malt. Holotype:UF25129, collected 25 June, 1971 by Jerry J. Landye. Paratypes: UF 358375 (7); ITCV (3); same data as the holotype.
Distribution. This species is known from the foothills of the Sierra La Catarina, which lies about 20 km southwest of Buenaventura. The range is oriented north-south, and is about 60 km long and about 10 km wide near its middle
Remarks. The reproductive system differs remarkably from other species of Humboldtiana by its very short genital atrium, by its long, slender penis retractor muscle which forms a sheath around the lower third of the epiphallus, and by its very long epiphallus, which is about 5-6 times the length of the penis. The flagellum is short, being less than the length of the epiphallus. The dart glands are associated close to the dart-sacs, and the free vagina is very short
Etymology. The species name spectabile (f.), is from the Latin spectabilis (m.), meaning worth seeing, notable or remarkable. The name alludes to the stately appearance of the shell.
Species from the Sierra Victorino
These are large, strongly marked species that are characterized anatomically by a long verge, by having the penis retractor muscle form a narrow collar around
Table 5. Humboldtiana spectabile n. sp. Measurements in mm of the holotype (UF 25129) and six paratypes (UF 358375).
H_W_AW AH Whorls H/W AW/W AH/H AW/AH
Holotype 35.4 38.3 21.1 28.3 4.2 0.92 0.55 0.80 0.75
Paratypes 
Min. 30.6 35.1 19.2 24.1 4.0 0.86 0.52 0.74 0.66
Max. 35.3 37.2 21.1 26.6 4.1 0.96 0.55 0.82 0.80
Avg. 33.13 36.59 19.77 26.46 4.09 0.91 0.54 0.79 0.75
SD 1.59 0.99 0.64 1.63 0.06 0.03 0.01 0.03 0.04