LANKESTERIANA


roadcuts in tropical wet premontane forest to 900
m of elevation in the Caribbean watershed of the
Talamanca range.

COSTA RICAN MATERIAL STUDIED: Cartago: Turrialba,
Tayutic, carretera entire Tayutic y Jicotea, Platanillo,
laderas del Rio Platanillo, siguiendo la margen del rio,
9049'27.46"N 83033'13.29"W, 878 m, bosque muy
humedo premontano, 1 abril 2008, D. Bogarin 4201,
A. Russell & R. Samuel (JBL-Spirit).

 This species is easily recognized by the pink
flowers and the strongly ruffled purple lip with 7
crisped golden-yellow keels. The leaves are bright
green and smooth. Plants from Panama and cultivated
at Lankester Botanical Garden produce fully opened
flowers. Nevertheless, the plant collected in Costa
Rica has flowers self-pollinated and developing fruits.
This condition has been also observed in Nicaraguan
material (Dressler, pers. comm. 2008). Sobralia
triandra A.H.Heller & A.D.Hawkes described from
Nicaragua could be conspecific with S. bouchei.

14. Warmingia zamorana Dodson, Icon. P1. Trop., II,
 6: t. 599. 1989. Type: Ecuador. Zamora-Chinchipe:
 Zamora, 1000 m, August 1968, C. H. Dodson
 3842 holotypee: SEL). Warmingia margaritacea
 B. Johans., Lindleyana 7: 194. 1992, syn. nov.
 Type: Costa Rica. [Cartago]: Turrialba, C.A.T.I.E.,
 epiphytic onHibiscus sp. forming hedge, 600 m, 19
 November 1988, B. Johansen & M. Sorensen 138
 holotypee: C, not seen).

DISTRIBUTION: Costa Rica and Ecuador.

ETYMOLOGY: named from the city of Zamora in southern
Ecuador, the locality of the type specimen.

HABITAT IN COSTA RICA: epiphytic in premontane wet
forest on garden trees and bushes in Turrialba region.

MATERIAL STUDIED. COSTA RICA. Cartago: Turrialba,
frente al edificio principal del Centro Agron6mico
Tropical de Investigaci6n y Ensefianza (CATIE) en
cerca viva a la par de un tubo de agua, 2 a 3 flores
pendientes, 3 noviembre 2003, A. Karremans 452.
(JBL-spirit) (Fig 13, 15-K). Turrialba, CATIE. Frente
al Edificio Principal del CATIE, sobre una cerca viva
de Hibiscus sp., al lado de un tubo de agua. 9053'22"


LANKESTERIANA8(2), August 2008. 0 Universidad de Costa Rica, 2008.


N 8339'12" W, 600 m, 4 noviembre 2005, A.
Karremans 1123 (JBL-spirit). ECUADOR. Zamora-
Chinchipe a lo largo del Rio Zamora entire 600 a 800
m, H. Medina s.n. (CIOA). Morona-Santiago: cerca de
Patuca, H. Medina s.n. (CIOA-spirit) (Fig. 15-L).

 This species was first recorded from Costa Rica
in 1992 by Johansen as a new species named W
margaritacea. The plant was collected at CATIE
campus in Turrialba, Costa Rica, growing in Hibiscus
sp. fences (Johansen 1992). After its description,
the species had been long known only by the type
collection (Atwood 1999, Dressler 2003). Fifteen years
later, in November 2003, botanical exploration carried
out at CATIE revealed more populations of this species
growing in Hibiscus sp. (Pupulin 2004, 2005).
 With a careful analysis of the type specimen of W
zamorana as well as of Ecuadorian material cultivated
from the Zamora-Chinchipe region, we conclude that
the characters used to separate W margaritacea from W
zamorana are inconsistent, and both species should be
considered conspecific. Johansen (1992) stated that W
margaritacea can be distinguished from W zamorana
by the coriaceous, lanceolate, subfalcate leaves, the
pauciflorus inflorescence, the rhombic petals, the rigid
column appendices and the self-pollinated flowers.
However, this set of features were also observed
in living specimens of W zamorana from Ecuador.
Morevover, Johansen stated that the flowers of W
margaritacea are pure ivory and pearly appearance,
contrasting W zamorana as having a yellow lip callus
(Dodson 1989), but the callus of all the examined Costa
Rican records is bright yellow in color (a photograph
is given in Pupulin 2005). The serrate petals and the
three-lobed lip with a bilobate yellow callus were
also observed in both Ecuadorian specimens from
Zamora and Costa Rican material collected at CATIE.
Although we have still not found a wild population of
this species outside of CATIE, some plants have been
collected growing in Cupressus lusitanicus trees.
 Warmingia zamorana has not been recorded
yet in the orchid floras of Panama and Colombia,
the two countries in the middle of its distribution.
Nevertheless, that disjunction may be attributed to
undercollection of these small and easily overlooked
epiphytes. It is remarkable that similar examples
of disjunction in the orchid floras of Costa Rica