LANKESTERIANA


 t. 26. 1838.
Pescatorea white (Rolfe) Dressier, comb. nov.
 Basionym: Zygopetalum white Rolfe, Gard. Chron.
 ser. 3, 7: 354. 1890.

Warczewiczella These species have been lumped
with Cochleanthes by some workers (Schultes &
Garay 1959), but the molecular data strongly support
the separation of Warczewiczella from Cochleanthes
as proposed by Fowlie (1969) together with some
more recently described species, but excluding W.
picta (here placed in Stenotyla). As defined here, the
genus consists of 10-12 species. This clade consists of
plants with relatively large flowers with a lip enfolding
the base of the column and with a thick, sulcate basal
callus composed of several or many ridges. The stipe
is shield-shaped. The lateral sepals are variable and are
rolled and backswept to form false spurs in some taxa
(e.g., W amazonica). The clade is well supported in
the combined tree with the exception of W wailesiana,
which is on a relatively long branch that is unsupported
as sister to the other Warczewiczella species in the
strict consensus of the shortest trees. Morphologically,
it agrees closely with other species in this clade, so
we include it in this genus even though it lacks strong
molecular support. The molecular data do not unite it
with any other clade, and we wish to avoid creation
of monotypic genera lacking strong molecular or
morphological support.

Type species: Warczewiczella discolor (Lindl.) Rchb.
f., Bot. Zeit. Berlin 10: 636. 1852.
= Warrea discolor Lindl., J. Hort. Soc. London 4: 265.
1849.
Warczewiczella guianensis (Lafontaine, Gerlach &
Senghas) Dressier, comb. nov.
 Basionym: Cochleanthes guianensis Lafontaine,
 Gerlach & Senghas, Die Orchidee 42: 285. 1991.
 Warczewiczella lobata (Garay) Dressier, comb. nov.
 Basionym: Cochleanthes lobata Garay, Orquideo-
 logia 4: 21. 1969.
 Warczewiczellapalatina (Senghas) Dressier, comb. nov.
 Basionym: Cochleanthes palatina Senghas, Die
 Orchidee 41: 96. 1990.

Stenotyla Dressier, gen. nov.
 Chondrorhynchae Lindley similis, sed mento
brevi, labelli callo basali, angusto, et pseudobulbis
manifesto evolutis lrI, i, i


 Type species: Chondrorhyncha lendyana Rchb.f.
Gard. Chron., n.s. 26: 103. 1886.
 Etymology: From the Greek stenos (narrow) and
tylo (callus), describing the shape of the lip callus.

The three species of this group possess a vegetative
similarity to Chondrorhyncha, but with small, distinct
pseudobulbs concealed in the leaf bases. Members of
this genus are clearly distinguished by the very narrow
2- or 4-toothed basal callus, the short chin and the
presence ofpseudobulbs. Pupulin (2000: 22) illustrates
S. lankesteriana.

Stenotyla lankesteriana (Pupulin) Dressier, comb. nov.
 Basionym: Chondrorhyncha lankesteriana Pupulin,
 Lindleyana 15: 21. 2000.
Stenotyla lendyana (Rchb.f.) Dressier, comb. nov.
 Basionym: Chondrorhyncha lendyana Rchb.f., Gard.
 Chron. n.s. 26: 103. 1886.
Stenotylapicta (Rchb.f.) Dressier, comb. nov.
 Basionym: Warczewiczella picta Rchb.f., Gard.
 Chron. n.s. 20: 8. 1883.

Chondrorhyncha helleri L.O.Williams is closely allied
to S. lendyana, but the material available is not sufficient
to show whether or not it is distinct. Chondrorhyncha
estrellensis Ames, if distinct, would also be included
in Stenotyla.

Cochleanthes With the recognition of Warczewic-
zella, Cochleanthes now consists of only two species:
the type, C. flabelliformis, and C. aromatica. The lip
does not enfold the base of the column, and the column
is distinctly keeled.

Although our sampling within the pseudobulbous
Zygopetalum grade (Fig. 5) is sparse, the cladograms
reveal several inconsistencies with current generic
concepts. Koellensteinia is polyphyletic, and the three
representative species are scattered within a clade
containing Acacallis, Otostylis, Paradisanthus, and
Zygosepalum. Zygopetalum is also polyphyletic; Z.
maxillare is sister to a clade containing Neogardneria
murrayana, Pabstia, and several other Zygopetalum
species. Finally, a strongly supported (100% bs) clade
contains Galeottia, Batemannia, and Zygosepalum
labiosum. Three of the Galeottia species plus
Batemannia lepida are on a very long branch relative
to the other members of this clade. As noted above, this
long branch is due to unusual (possibly paralogous)


Vol. 5, N 2