WHTTEN ET AL. Relationships of Zygopetalinae


does not fit any other group now known, though
superficially similar to Chondroscaphe. The lip has
a strong median keel basal to the two-lobed callus,
the blade of the lip is pubescent, and the viscidium is
truncate and subtriangular. The term rhynchos (snout
or muzzle) is intended more to suggest a relationship
to Chondrorhyncha than to describe a specific feature
of the flower. The genus consists of a single known
species. Drawings are given by Jenny (1989: 93).

Aetheorhyncha andreettae (Jenny) Dressier, comb. nov.
 Basionym: Chondrorhyncha andreettae Jenny, Die
 Orchidee 40(3): 92. 1989.

Ixyophora Dressier, gen. nov.
 Warczewiczellae Rchb.f similis, sed stipite prope
viscidium panduriformi vel angusto, I I, 1 i
 Type species: Chondrorhynchaviridisepala Senghas,
Die Orchidee 40(5): 181, f. 1989.
 Fi n1 .1. *r From the Greek ixys, waist, andphoreus,
bearer or carrier, referring to the narrow "waist" of the
stipe.

 Superficially similar to Warczewiczella, these species
are distinctive in the form of the stipe that is narrowed
between the viscidium and the pollinaria. These
species form a group sister to Chaubardiella (Fig. 4).
Senghas (1989: 180) gives a drawing of L viridisepala
and Senghas and Gerlach (1991: 283) illustrate L.
aurantiaca.

Ixyophora aurantiaca (Senghas & Gerlach) Dressier,
comb. nov.
 Basionym: Chondrorhyncha aurantiaca Senghas &
 Gerlach, Die Orchidee 42: 282. 1991.
Ixyophora carinata (Ortiz) Dressier, comb. nov.
 Basionym: Chondrorhyncha carinata Ortiz, Orqui-
 deologia 19(2): 18. 1994.
Ixyophora viridisepala (Senghas) Dressier, comb. nov.
 Basionym: Chondrorhyncha viridisepala Senghas,
 Die Orchidee 40: 181. 1989.

 Chaubardiella Garay This uniform group (about
eight species) has nonresupinate flowers and deeply
concave lips; the viscidia are placed on the legs of its
pollinators. The combined analysis forms a strongly
supported group (Fig. 4).

Pescatorea (16 spp.) and Bollea (12 spp.) have long
been regarded as sister taxa. As traditionally defined,
these genera differ primarily in the relative width of
the column (much broader in Bollea), but possess


similar floral shapes and are known to form natural
"intergeneric" hybrids. In our sampling, the four
species of Pescatorea are intercalated among the three
representatives of Bollea on short branches; together,
the clade is well supported. Given these relatively trivial
morphological distinctions and the lack of molecular
support for maintaining them as separate genera, we
conclude they should be treated as a single genus. Both
genera were described in the same publication, giving
neither name clear priority. Since Bollea contains fewer
species than Pescatorea, we choose Pescatorea as the
generic name for this clade to minimize the number
of nomenclatural transfers. The name Pescatorea is
adopted here and the originalPescatoria is considered a
typographical or orthographical error, the generic name
being derived from the personal name Pescatore. The
name Pescatoria was not an intentional latinization, as
the author himself adopted the alternative spelling in
successive publications. The correction does not affect
the first syllable of the name or the first letter of the
name. The generic name Pescatorea is formed with the
addition of the letter "a" to the customary spelling of the
personal name, as recommended by the International
Code of Botanical Nomenclature (ICBN).

Pescatorea Rchb.f, Bot. Zeitung (Berlin) 10: 667.
1852. [as Pescatoria, orth. var.]
Type species: Pescatorea cerina (Lindl. & Paxton)
Rchb.f, Bot. Zeitung (Berlin) 10: 667. 1852.
Pescatorea coelestis (Rchb.f.) Dressier, comb. nov.
 Basionym: Bollea coelestis Rchb.f, Gard. Chron.
 ser. 2, 5: 756. 1876.
Pescatorea ecuadorana (Dodson) Dressier, comb. nov.
 Basionym: Bollea ecuadorana Dodson, Selbyana 7:
 354. 1984.
Pescatorea hemixantha (Rchb.f.) Dressier, comb. nov.
 Basionym: Bollea hemixantha Rchb.f, Gard. Chron.
 ser. 3, 4: 206. 1888.
Pescatorea hirtzii (Waldvogel) Dressier, comb. nov.
 Basionym: Bollea hirtzii Waldvogel, Die Orchidee
 33(4): 143. 1982
Pescatorea lalindei (Linden) Dressier, comb. nov.
 Basionym: Batemannia lalindei Linden, Numer. List
 90. 1873.
Pescatorea lawrenceana (Rchb.f.) Dressier, comb. nov.
 Basionym: Bollea lawrenceana Rchb.f, Gard.
 Chron. ser. 2, 15: 462. 1881.
Pescatoreapulvinaris (Rchb.f.) Dressier, comb. nov.
 Basionym: Bollea pulvinaris Rchb.f, Linnaea 41:
 107. 1877.
Pescatorea violacea (Lindl.) Dressier, comb. nov.
 Basionym: Huntleya violacea Lindl., Sert. Orchid.


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