During times of extremely low tides, however, we have observed the tops of the Acropora reef protruding from the water, exposure of the Porites-Penicillus back-reef flats, and apparent cessation of water movement in the Lagoon. Both McNulty (1968) and Michel (1970) also observed simi- lar periodic loss of circulation. At such times, with calm weather, the southerly movement of Lagoon water is unopposed by the usual wind driven waves from the reefs, and hyper- saline or storm water may move south from the northwest Lagoon into the channels around Patricia and Bovoni Cays. This rather unique interaction of waves and tides and its effects on circulation and flushing of the Lagoon is crucial to the hydrography and ecology of the area. One of the most obvious of these effects is that increased circulation causes the water in Benner Bay and the Lagoon to become more homogenous, generally reflected by a reduc- tion of the usually higher salinities in northeast Benner Bay and northwest Lagoon to levels closer to those of the open sea to the south. This effect is less marked at deeper more exposed stations (2, 4, 11) which normally maintain salini- ties close to open sea values, and, as a consequence, the range of salinity throughout the area is compressed. On the other hand, during periods of poor circulation evaporation increases salinity in the more remote areas and distinct spatial heterogeneity develops. Page 23