BONACCORSO: A PANAMANIAN BAT COMMUNITY During the March-May sampling, 27 percent of the A. jamaicensis adult females captured were nulliparous (Fig. 12,), and nearly all of these probably had been born in the first or second birth pulse of the previous year (8 to 12 months before). Through the next three bi- monthly periods the percentage of nulliparous females in the popula- tion steadily rose to 49 percent, as recruitment occurred from the off- spring of the year. Between November and the following March the proportion of nulliparous females declined, not from their having given birth, but presumably from higher mortality and/or dispersal rates than are found in older females. GROUNDSTORY FRUGIVORE GUILD Groundstory frugivores follow a reproductive pattern similar to canopy frugivores. There are two births per year per female, one in the dry-to-wet season transition and the other in mid-wet season (Fig. 14). Simultaneous pregnancy and lactation in March-May and recaptures of marked individuals indicate that females have a postpartum estrus. Females are reproductively inactive from mid-October through December. Artibeus phaeotis 1001 Pregnant 90- j Lactating 80 r Nonreproducing V) S-70 60 o 50 O 40 z U 30 20- 10- few data J-M M-M M-J J-S S-N N-D N=28 14 14 15 19 FIGURE 13.-Reproductive timing in female Artibeus phaeotis. 1979