280 TELFORD: MULTIMAMMATE RAT POPULATION BIOLOGY IN TANZANIA samples (Tables 13, 14) demonstrated that mean HBL and mean weight of female P. natalensis were greater in the critical December-February and March-April periods of 1983-84 than at comparable times in 1981-82 and 1982- 83. This probably reflects an increased food supply despite greater population density during the exceptionally heavy rainfall of December 1983-January 1984. The data also could indicate survival of the larger members of the cohort, with smaller individuals having greater mortality or being forced to emigrate from the study areas. CONCLUSIONS The data obtained during this study suggest the following conclusions: 1) In dry years, reproduction in either uncultivated or cultivated habitats begins in April and ends in August. Populations begin reproduction in those years at minimal density levels. REPROD. I POST-REP. El NON-REP. D 1982 1983 1984 1985 Figure 26. Structure of the female cohort from the university campus P. natalensis population in fallow maize field or uncultivated grass microhabitats in January of each year, 1982-85. TABLE 12. Average trap success forPraomys natalensis by habitat and season. % Positive traps in Season Maize fields Uncultivated grass July-December 31.2 17.1 January-April 12.7 7.3 May-June 4.0 5.4