WILLIAMS: CHASMODES TAXONOMY


FIGURE 6.-Holotype of Chasmodes bosquianus longimaxilla new subspecies. Adult male, 86
mm SL, from Dauphin Island, Alabama (USNM 219830).

Table4); like C. b. bosquianus, number of mandibular teeth increases with
increased size (Fig. 4); pores in both mandibular series combined 4 to 5
(rarely 6; X = 4.1, n = 115; Table 1). Of the three taxa referable to Chas-
modes, C. b. longimaxilla alone shows a difference in number of small
upper pharyngeal teeth between size classes (Table 1). Enlarged upper
pharyngeal teeth 5-7 on each side (Table 1); each lower pharyngeal with 4-5
small teeth anterior to 4 large, recurved canines; maximum size approxi-
mately 85 mm SL.
 Pigmentation and other characters of larvae immediately after hatching
are similar to those described by Hildebrand and Cable (1938) for C.
bosquianus (= C. b. bosquianus) from Chesapeake Bay.
 The only small (<10 mm) posthatching, wild-caught specimen of C. b.
longimaxilla examined measures 8.9 mm SL, and has the following charac-
ters: maxillary length 11% SL, 11 mandibular teeth, 4 pores in mandibular
series, and no developed lip flaps on lower jaw. The 5 to 7 preopercular
spines present in larval specimens of C. b. bosquianus are represented by 5
serrations in this specimen.
 Pigmentation of this small individual is consistent with that described
previously for adult female Chasmodes. All young of this subspecies, as
well as other members of the genus, have the mottled pigmentation pattern
until they attain a length of 35 to 40 mm SL, at which time the males begin to
show the characteristic adult color pattern (described previously).
 STATISTICAL COMPARISONS.-Statistical comparisons of populations of
Chasmodes bosquianus longimaxilla with populations of C. saburrae and
C. b. bosquianus are discussed in the generic and the other subspecies
accounts, respectively. A statistically significant difference in comparison
of regression equations (Table 5) for maxillary length and number of
mandibular teeth between populations of C. b. longimaxilla was found in
only one comparison (Tables 6, 7). A significant difference in maxillary