BULLETIN FLORIDA STATE MUSEUM


 I thank the curators and assistant curators of the fish collections from which material was
borrowed. Special appreciation is accorded Victor G. Springer for providing access to
specimens in the United States National Museum of Natural History.
 I wish to express special thanks to R. Douglas Nester for his assistance in collecting
specimens and for his optimism and encouragement regarding my research. Thanks are also
accorded Douglas Clarke for providing the holotype of the new subspecies. Carter R. Gilbert,
George H. Burgess, and Victor G. Springer critically reviewed and improved earlier versions
of the manuscript. Grace Russell typed and helped edit the manuscript.
 I thank my parents for their support and encouragement. Special consideration is due my
wife Karen for her understanding and support throughout this study.

 METHODS
 Measurements were made with dial calipers and recorded to the nearest 0.1 mm. All counts
and most measurements were made using a Wild M5 stereoscope. Meristic counts and
morphometric measurements follow Hubbs and Lagler (1958) and Smith-Vaniz (1976).
Maxillary length was measured from the tip of the snout to the posterior edge of the left
maxilla. The edge was occluded from view by an opaque covering of skin. The posterior edge
was located with a sharp probe or jewelers forceps. Counts of pores in the mandibular series
(Fig. 1A, B) include all pores anterior to the narrow space between the preopercular and
articular bones on each side of the head (Fig. 1). This space can usually be seen as a narrow
translucent area immediately ventrad to the posterior edge of the maxilla. Caudal ray counts
include all segmented rays. The branchiostegal membrane is broadly fused to the body and
was cut on the right side to provide access to the branchial cavity. Counts of gill rakers and
pseudobranchial filaments refer to those on the right side of the body. Gill raker counts
include those rakers on the upper and lower limbs of the first arch (no rudiments were
observed). Upper pharyngeal teeth are located by separating the second and third gill arches
with jewelers forceps and, using illumination from beneath the specimen, are found in two
small semicircular clusters on the posterodorsal roof of the branchial cavity. As the lower
pharyngeal teeth must be dissected out of the specimen to be counted, my counts are based on
examination of four specimens of each subspecies of Chasmodes bosquianus and eight
specimens of C. saburrae. Vertebral counts were taken from radiographs and cleared and
stained specimens. Precaudal vertebrae included vertebrae lacking a well-developed hemal
spine. Caudal vertebrae include vertebrae with a well-developed hemal spine; posteriormost
caudal vertebra bears the dorsal and ventral hypural plates.
 Information regarding embryonic development and posthatching characters was obtained
from eggs and larvae taken from experimental laboratory aquaria.
 Numbers in parentheses in material examined sections refer to the number of specimens in
that particular lot.
 Linear regression equations and their correlation coefficients, and covariance comparisons
(Sokal and Rohlf 1969) were computed using a covariance program written by Dr. S. A.
Bloom (Zoology Department, University of Florida). Computations were made utilizing the
facilities of the Northeast Regional Data Center of the State University System of Florida,
located on the campus of the University of Florida. F values are not included for covariance
comparisons presented in matrix form, but are available upon request.
 Because of editorial problems, all the populations of each form are not differentiated in
Figs. 3 and 4. Maxillary length as percent of SL is presented in tabular form for selected
populations (Table 8). Graphs of the number of mandibular teeth vs. SL were prepared for
each population to facilitate comparisons but are not included owing to space limitations.
 Repositories, with abbreviations in parentheses, of specimens examined are: Academy of
Natural Sciences of Philadelphia (ANSP), British Museum (Natural History) (BMNH), Cali-
fornia Academy of Sciences (CAS-SU), Field Museum of Natural History (FMNH), Florida
Department of Natural Resources, Marine Research Laboratory, St. Petersburg (FSBC),


VOL. 29, NO. 2