WILLIAMS: CHASMODES TAXONOMY


comprehensive taxonomic treatment of the genus is that of Springer (1959),
who recognized two species, C. bosquianus and C. saburrae. Bath (1977)
assigned eastern Pacific species (C. jenkinsi and C. maculipinna) to Hyp-
soblennius. Springer and Gomon (1975) placed C. herklotsi in the synon-
ymy of Omobranchus fasciolatoceps.
 Norman (1943) provided a synopsis of the genera in the family Blennii-
dae. Included were three species in the genus Chasmodes: C. bosquianus,
C. novemlineatus, and C. maculipinna. He apparently overlooked C.
saburrae. Springer (1959) placed C. novemlineatus in the synonymy of C.
bosquianus.
 In an osteological analysis of the family Blenniidae, Spring (1968)
mentioned Chasmodes several times, but only in regard to intergeneric
relationships.
 Bath (1977), in his review of the tribe Blenniini, analyzed phylogenetic
relationships within the group. He suggested that Chasmodes serves as a
link between the generaHypsoblennius andHypleurochilus, but he did not
examine the relationship between the two species of Chasmodes.
 In a review of the western Atlantic species of Hypsoblennius, Smith-
Vaniz (1980) presented characters to distinguish Chasmodes from Hypso-
blennius and Hypleurochilus.
 Ecological studies dealing with Chasmodes saburrae appear in Reid
(1954) and Springer and Woodburn (1960). Tavolga (1958) presented his
results of studies on underwater sounds produced by specimens of C.
saburrae (erroneously called C. bosquianus). Phillips (1971a, 1971b, 1974,
1977) reported on the social behavior and utilization of space by C.
bosquianus.
 Hildebrand and Cable (1938) presented information on the develop-
ment and life history of Chasmodes bosquianus.
 Additional references to the genus Chasmodes are in the form of geo-
graphical records and/or ecological notes (Weymouth [1910] for Louisi-
ana; Baughman [1950] and Hoese [1958] for Texas; Tabb and Manning
[1961] for Whitewater Bay, Florida; and Ogren and Brusher [1977] for
northwest Florida).
 In this paper, Chasmodes bosquianus (Lacepede 1800) and C. saburrae
Jordan and Gilbert 1883 are considered valid species, and a new subspe-
cies, C. b. longimaxilla, is described from the northern Gulf of Mexico.
 ACKNOWLEDGEMENTS
 I wish to express my appreciation to Robert L. Shipp, who provided encouragement and
guidance during this research and also arranged for financial assistance. I thank Joseph F.
Fitzpatrick, Jr., and Stephen A. Bloom for assistance with statistical analysis of the data; Alvin
P. Rainosek for consultation on statistical procedures; and James M. Boyles, David M. Dean,
and George F. Crozier, for advice and suggestions regarding my research.
 Chris Benedict kindly provided the material that Kerry MacPherson of North Carolina
Power and Light Company donated to the Florida State Museum.