BULLETIN FLORIDA MUSEUM NATURAL HISTORY VOL. 38 PT. I (3) TEMPORARY PONDS Defining a Temporary Pond Temporary wetlands can be defined as natural bodies of water which experience a recurrent dry phase of varying duration (Williams 1987). The hydrologic emphasis in these wetlands is on the cyclic nature of drying and re-filling as permanent waterbodies are also capable of drying completely in exceptional years. In addition, selection has occurred in temporary wetlands for species adapted to this cyclic drying and filling and results in periodic bursts of productivity that fuel these systems (Patrick and Khalid 1974; Brinson et al. 1981; Reddy and Graetz 1988). A temporary pond is a small (generally less than 5 hectares), isolated, temporary wetland that is depressional in nature. There is considerable confusion of terms in dealing with these small temporary wetlands, both among regulatory agencies and in the literature. Terms used to describe them include flatwoods marshes/ponds, ephemeral ponds/wetlands, highlands marshes, pineland depressions, depression meadows/marshes, St. John's wort ponds, seasonal marshes/ponds, intermittent ponds, and vernal pools (Holland 1988; Florida Natural Areas Inventory and Florida Department of Natural Resources 1990; Kushlan 1990). Some of these terms allude to the fact that temporary ponds are difficult to identify as wetlands during their dry cycle (Means 1990). Hydrology Hydrology is the major factor influencing and maintaining the community of wetland plants in the temporary pond basin (Mitsch and Gosselink 1986). External water inputs transport nutrients into the system and mobilize those bound in vegetation and soil. Water depth, duration, and frequency of flooding all influence the formation of hydric soils and the presence of hydrophytes within the pond (Gosselink and Turner 1978). The hydroperiod of temporary ponds is highly variable, depending upon elevation, basin characteristics, and rainfall patterns (Means 1990). Some ponds may fill and dry on an annual basis while others may contain water only in the wettest years. When a temporary pond drains, productivity is lowered and aerobic decomposers begin rapidly breaking down the accumulated organic matter (Cole and Fisher 1979; Reddy et al. 1986). Temporary ponds are detrital systems, and most of the organic production decomposes before entering the detrital food chain (Mitsch and Gosselink 1986). The vegetation of the pond basin is readily and almost completely decomposed on a cyclic basis (Gopal 1986). The breakdown of