270 The cumulative acreage of water coverage under primitive conditions is greater than double that under present conditions in wet years, but there is less difference in cumulative acreage between the two conditions in dry years. When a series of three or four dry years occurs, there is greater cumulative water coverage under primitive conditions than under present conditions for the first few years, but about the same regional-scale hydroperiod under the two conditions for the last year or two of the drought (Figure 63). This is apparent in the graph for the years 1954-55, 1955-56, and 1956-57 and for the years 1963-64, 1964-65, and 1965-66. Fluctuation between extremes in wet years and dry years is greater under primitive than under present conditions. Figure 64 indicates the net annual change in water in storage in the soil and substrate and in the ponds, sloughs, and strands under present and primitive conditions as simulated by the model. Water in storage is the difference between the total rain received in a year and the total water that leaves the system through runoff and ET. Although values for the two parameters, runoff and ET, fluctuate more from one year to the next under present conditions than under primitive conditions, change in storage is just the opposite. Change in storage fluctuates less under present conditions. The natural system, without drainage, is able both to store more water and to release