74 Equation for surface water runoff. Runoff in the water model is governed by the Manning equation (Lindsley et al., 1958): = 1.49 hw 5/3 L (sin 0)1/2 (11) where QW is flow rate, hwis depth of the surface water, L is the width of overland flow, and 8 is h/d, or the land elevation gradient between upper and lower ends of the flow-way. One quarter of the water depth at the deepest local point (from the curve relating depth at deepest local point to total surface water volume) was the estimate of h used in the model. Runoff values simulated with the use of this estimate of h were very similar to those simulated with the use of "average depth," a value determined by dividing total water volume by total area of surface water. The first estimate, however, yielded a more realistic simulation of seasonal change in surface water area. The equation is: hw= c h, (12) where h' is depth at local deepest point and c is 0.25. In the model L was a variable that changed according to a power of the proportion of the total wetlands area containing surface water. The equation is: