Figure 3 shows the long term mean monthly moisture relations in the vicinity of the study area (Royal Palm Ranger Station, in the Everglades near the Tamiami Trail, latitude 25000'N). The graph demonstrates an excess of rainfall over evaporation during the summer and early fall, but a moisture deficit, evaporation greater than rainfall, during the spring and summer. In south Florida, pan evaporation appears closely related to solar radiation, also shown in Figure 3. Transpiration, the loss of water through plants, changes seasonally. Transpiration rates for some plants, such as cypress, willow, and many marsh macrophytes are greatly reduced when the plants shed their leaves or die back in late fall, suspending photosynthesis during winter. The Study Region The study region can be roughly defined as the noncoastal parts of Lee, Hendry, and Collier counties. It also includes the seasonally drying marshes on the western and northwestern shores of Lake Okeechobee and parts of Charlotte and Glades counties (Figure 4). This area, which lies between latitudes 250 47' and 270 07' N, comprises numerous sloughs, marshes, cypress strands, wet prairies, and ponds that are linked by flows of energy through the movement of water and animals such as wading birds. Although many of these wetlands are in their natural state,