Breeding takes place only when average bird weight rises above a threshold value. In the model, breeding success of one species, the Wood Stork is taken as an index not only of the condition of the area's wading bird populations but also of the general healthy functioning and overall productivity of the wetlands system. The model tests the effect of seasonal and long-term variations in rainfall and the effect of drainage on expansion and contraction of surface water area, fluctuation of the water table, total annual fish production, fish density, and breeding success of Wood Storks. Climate Figure 2 reveals seasonal and long-term patterns in the rainfall of southwest Florida. The record of rainfall is for the years 1951 through 1975 collected by ALICO (Atlantic Land Improvement Company, La Belle) at Corkscrew grove, a commercial orchard at latitude 260 25'N, located 30 km from the coast and immediately north of Corkscrew Swamp (see Figure 4 for location). 5. Average yearly rainfall was 1421 mm at Corkscrew grove, which is slightly greater than the 1372 mm average reported by the U. S. Weather Bureau near the Gulf Coast at Page Field in Ft. Myers, Lee County. On the average, Corkscrew grove received 68% of its annual rain during the 5 mos from June through October.