There are approximately 30,000 ponds ranging in size from less than an acre to more than 100 acres in Lee, Hendry, and Collier counties. Freshwater wetlands once comprised 53% of the total area. Drainage, which has accelerated in the past 15 yrs, has reduced wetlands area by more than 60?, diminishing the capacity of the area to store water from wet years. Regional-scale depth-area, area-volume, and depth- volume curves were developed to relate the intensity and timing of rain to the seasonally changing area of surface water. Classical hydrologic equations were combined with the regional curves to produce a water model simulating seasonal and long-term water storage and runoff. Patterns of land area covered by water from the water model were used to drive three models relating water patterns to fish and storks. Seasonal oscillation of water area increases the efficiency of the flow of energy from prey to predator and allows a system to support more top consumers than would otherwise be possible. Concentration of the fish production of the wetlands of southwest Florida is increased approximately 25 times by the sun's energy, acting seasonally through evaporation and the transpiration of plants to shrink water area. This extra concentration factor in the food chain gives a very high "energy quality" value to the Wood Stork, which may serve as an Indicator of the productivity of wetlands of the region, viii