10 Biological Theory The need to consider the biological characteristics of fish popula- tions in economic analyses of fisheries becomes immediately evident when the nature of the resource (fish population) is considered. Fish popu- lations can be placed in the category of use-dependent flow resources with a critical zone (Schaefer, 1957). A critical zone as used here is defined to be a rate of decrease in flow which cannot be reversed economically or technologically. Thus, in terms of fish population, a critical zone would correspond to that level of population which has an insufficient reproductive potential to remain viable. Sustainable Yield The main aspect of biological theory that is relevant to production analysis of a fishery is that of population dynamics. More precisely, the notion of sustainable yield (SY) is one of the cornerstones of bio- economic theory. The size of a non-exploited fish population (biomass) can, in general, be assumed to be a function of three factors: growth, recruit- ment and natural mortality (Gulland, 1965). Each of these variables, in turn, is a function of the biomass. Individual growth is generally assumed to be at a maximum at low levels of population and to decrease as the population size increases. Natural mortality acts in the oppo- site manner, being low at low levels of population. As, population size increases, natural mortality increases due to increased competition for food and other such factors. Recruitment, the rate at which individuals enter into the fishable population, is generally assumed to be low at